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A chicken embryo gene expression database

	Anatomical Location
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	Gene Name
	GEISHA ID
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VCAN
versican

The human orthologue of this gene is associated
with the following human disease:

Wagner syndrome 1

Gene

Official Symbol	VCAN
Official Full Name	versican
CGNC ID	49376
also known as	PG-M, versican core protein\|chondroitin sulfate proteoglycan 2\|chondroitin sulfate proteoglycan core protein 2\|large fibroblast proteoglycan\|proteoglycan
gene type	protein-coding

Genomic Map

	Ensembl ID not known
	NCBI
	UCSC

Sequence Information

Genomic
RNA	NM_204787
Polypeptide	NP_990118

Gene Expression

In Situ Hybridization

GEISHA

Orthology

	Entrez Gene	Ensembl Gene	Genetic Phenotypes	MOD
Fruit Fly
Human	1462	ENSG00000038427	MIM:143200	MIM:118661
Mouse	13003	ENSMUSG00000021614	All phenotypic alleles (3):Targeted, knock-out(1) Targeted, other(1) Transgenic(1)	MGI:102889
Xenopus	100101672, 100037127			922301
Zebrafish	116993, 323465	ENSDARG00000009401	ZDB-GENE-011023-1, ZDB-GENE-030131-2185	ZFIN:ZDB-GENE-011023-1, ZFIN:ZDB-GENE-030131-2185

Gene Ontology

Molecular Function	calcium ion binding, carbohydrate binding, hyaluronic acid binding
Biological Process	cell adhesion
Cellular Component	proteinaceous extracellular matrix

Links to other databases

Entrez Gene	395565
Ensembl Gene
KEGG	gga:395565

Entry 1

GEISHA Id	VERSICANV3.Hudson.2010	Data Source	Publication
Complete cDNA Template Probe	show TACCTCAGACTGTCACAGATGGTGAAATAAGCCCTTATGATACGCTGGGAAGGACTGAATATGATGTTTCACCTAGGTT AACAGAGAGCACATCTGCAGCTTTGGAAGTGGAGCACACTTACTCTGAAGCAGAATTGTCCGAGGAACAAGGTAGGTCT GAAAGCACTGAGGACGCTTTCTTGACCTCTGTAGTTTTTCAGGATAGCACAGCTGTAGCTAAGAGTTCCACTGGTTCAT GGGAAGATATTGAAACAGGAGATACACAAAAACATGATGGTGATAATCAGACTGAACAAATAGAAGTGGGTCCTGTGAT GACAGCTACAGATAGCTTGGTACCAGCTTCTCAGAGAGAGTTGCCCAGGACAGGGTCTTCAGTTTCACTAACAAAAGAA AATCTGTATCTTGGTTCTCACTCAACAAAAGAACCCACAAAAAAATCAATGGAAGCAAAATCTGACAAGAAACTTACAA CTGTTGTAATCCCTAAGGCTTTGTTCACTGATCAGTATGATCTTACTACAGGGGGGGAGGGAAGAGAGAGCATGTACAC CGTTATGCCTGATAGAGTTTCTGGCGTGGCTCTGGTTAGCATCCCAGAATCAGATGTTCCTGCTGTATCAGAGACACTC ATGGATGAGCTTGCGGTAACCACTGGACAATCTTCTACAGCAGATGAGTCAACTCCATTCATTAAATTTAGTTCTTCTG CAACTGAGTTGGATAATGAGGCGTCAGCTGAGGGAAGCAGAGAGGACTTAAAAGATGTGCATCTCACCACGTCATCTGG AATACCTGTATCCTTTACTTTATTCACCGCCAATGAGACAGGATCTGAGGTCACTGCTCTCTCAGAAAGTACCAGTGCC CCACAGAAGTTTGAAGAAGGCATCACATCTGTTCTTCACTCATCGCAGCAAACAGAAGGATCGGCTATTTTAGAAAAGC AGGAAAAAACAAAGGAGCCAGAAATGAGTACAATAGATGCTAAAGTCCTTTATATCACAACTGTTGTTCCTGCTTCTGT TACGGCAGGATCTGAGGGACGCTTTGGAAGTGAGAAGTTCACGCACACTCCTCCAGTTTCTGGCATGTGGCTGCAGACA GATAAGGATCAGGTCTACATGACAGAAGAAACATCTCATACTAAGAGAATTGAGTTAGACACTGAAGATGATATCTCTG GAATGGAGCCTACATCATCTCCAGGGCAAATTATAGAATATACAAAGCATCTAGGAGCTCCAGTTTCAGCAGTAACAGA TGAAACCAAGACAAGCATGGAGACAGCAGAAACAGAAAGTGATGAAGAAGTAGTATCAGCTGATTTTGATCAGACTAAA GGTACTACAGAGGTCTTCCATACAAGCAGCTCTTTAGATTTGGAAAAGTTTACTCTATCCAAAATACCAGAGGATGAAA GTAGTGCAACAGTGAAATCGTTTAGCTCCTCAAGTGGTACTGTGTTACCAACTGCTGTGGCCACTGTCTTGGAGGTGAC TGATCATGAAGCAGATGAAACATCAGGATATGTTTTGAACATGACCTTTTCTACCCCAGAAGGTGAACAAAGAAAGGCT ACTGAGAAATCACCTGCAACTTCTGCTGAAGATGAAGTCTCTACAGGAACAGAGATCTCAAAATACACAATGACAGAAG GCGGCCAGATAAGCAGTGTGACATCTGCTGAGAAGGAGTCAGTGGCAGCACTTCAGGAGAGGGAAGAACAACCGTCTGT TGGCTTACCAGAAACAAAGGAACCCTTTAAGTTTACAGATGTAACAGAGATAGAGACTACAGTTCCACAGAGAGAAGGT GATACTTCCCTAGTTCCAGTCACTGTAGGTAGTGAGGACATTGGAGAGATGCAAGTTACTGACCATACTTCTTTTGACA GTATCATTCATACTGAAGCAACAGTAACAAGTACAAAAGCTAGTGAAGTGTTTCCTAAGGAGCTCTCCACAAAAGATCA AGATAGGGAACTAGGTACTGCCATGGGGTCTACCTTACCTGTGACATCTGTCCAAATGCATGAACAAAAGACGACAGCA GGATTTGAATCACCTCAAACAACCACTCAGGAAAAACATGACGAGATGGGTTCAGCCTATGATGAGATGTATCCAGCAA CAGAATTATCAGTGCCTGCATTAATGCTTACAGAATATGGACAAGTTTCAGGGCCTGTTGAAACTAGCACCAGGTCTTT GCATCTCACAGGGACTCCCAAAGCTGAGACTGCTACTGACCAAGAAGAGAAGATTACAGAAGCAGTGCCTGTAACATTT GGTACACAAGCAAAAGTGTATGAAAGCAAAGGAACCACCACAAGGGAAGAAGACAGAGATGTAGGAAGCTGGAATTCTG TGTTGCCCCCCCATACGATGCTGAGTAGTCCTTCTACTGCAGGAAGTATTAGTCTTCTGACTTTGGGAGCTTCTCCAAG TCAAACTCCAGAAGGTTCAGGAATATCAGAAGAGCTTGAAGAAGTCAAAACAGTTCCATTTTCATCTAGGGCAACAGAT AAGACAACTGTTATTAGTGATTTAACTACATCATCTATTAGCGCTGTAGACAAAATTCAGCCTACATCAGCATCCAAAC CTTTTGTTTCTTCAAAGTCGCCACGTATTATTCCTGAGGAAGATGAGGAGGTAACAAGCAGTGACATCATAGTAATTGA TGAATCTATTTCTCCAAGTAAAGCCAGTGCTGAAGATGATCTGACAGGAAAGATGGTAGAACCAGAAATTGATAAGGAA TATTTCACATCATCAACTGCTACTGCAGTTGCACGGCCTACTGCACCACCCACAGTGATGGAGGCCACAGAGGCCTTAC AACCACAAGAGGTGTCTCCTACTTCACACCCTGACAGTGGAACTGACATAAGATTGTATGTTATTCAAATCACAGGCAA TGATACAGATCATCCGGTGAATGAGTTTTTGGATCTGTTCAGTCGCCACATTCTTCCTCATGCAGTAGATGAGACCCAT ACTGATGCAGAATCAGCTCAGACTGAACCCTGTACTTCAGACTCTGTGCAGGATTCGTCTGAATATATAATATTGGATC CTTTCTTTCCAAACTTTATGGACTTTGAAGAGGAGGAAGAGGACTGTGAAAATACTACTGATGTTACAACTCCTCCAGC TTTGCAGTTCATCAATGGAAAGCAACAAGTGACAAGTGCACCTAAAAGCACTAAAGCTGAGGAAGCAAGAAGTGACCAA ATTGAAAGTGTTGCACATTCAAAAAATGTAACTTTTTCACAAATCAATGAAACAAACACCTTCATAATATCTGAAACAG AAGCTTCTGGTACAATGCAGCCAAGCAAAGCTGGAGAAGTAATGGGGGCATTTGAAGTCACACAGCCGACAGCAGATGT TGCAATGTTAGAACCAGTATATAGTGGTGAGTCAGAAGTTACCACAACAGACAAATATTTAGAGATTACGTCTGTGTAT GAGCAGTCACCAAAAAAAAATAAGGAGACTGTAATGTGGCATGGAACTGAGGAGAGCTCTACCAAAGACACAAAAAACT TGCTTTTGATCACTAATGAATCTTCTGGAGATGGCTCCACTGAATCTGATTTGTCTAGGTCTGTGTTCACAGAAATTTT GACAATGTCAAGTCATGAAGATAGTGAAAAAATTTCTCATACAACTTCTGTTCCTACTATCCTTTCTGTGGAAAGATCT GCAGTAACTGCTGCTCCGAGTGCAGATTCTGATACTGCTACTGTAGGCATAGATGTGAAAGACCTTATTCCAAAAGGTG GGACCGCTACTCCAGGAAATTACTATAAATCGACTATTAAACTTGATGCAGAATTCCCTTTTGAAAGCAACCCAGAAGC AACAAGTCATACAACAAAGCCTGACATGACTGCTTCTTCTTTTATAGTTCTGGAAGGTTCTGGAGATGTAGAAGAGAAC AGCACTTTAGCTTCAGCCATGACAACAGAAACAGCAGTTGCAGAAACACTTTCAGTGCAAGATACTTCTTTGGGTTCTG GCACAGTACTGCCAACAGAAATTTCTGTAACCATTTCAGAAATCACCCCTGCTTTGCCTGGAGGAACAAGAATTCTTTA TTCTACTTTTGATCAAAGTTCTGAAGCAACTGTTTCCACCAATTTTGTGTCTGAATTAATTATGGAGCAAGTAGTTGGC AGCTCTGTTGCAACTGAAAAGAAAGTTGAAGATGAAAAAGAAGTACAGACTACCGTTTATTCGAGTCAGGAAATTTCAA CCACTGATGCAAAAGGAAAATCAGAACTGGATGAGTTTGGAAGCACTACTAATGAAGTCAGAACTGTAAGTCAGGAACC CACTCCTCTCAGAGAAATAGTACCAATAACAGGTACCATGCATTCTGAAATCAAAAAGGTTACCGCCACTCCTTTCTTG AGAGAGAAGCTTTTTATAAATGAAGGATCAGCTGAAGAACCAGCAGACTTATTTGCAGGGAGTCCCACAAGAAAAGTGG TAAGTACTGACTCCCCATTTACTGATTCAGGTTCTGGAGACATAGATGTTATCACTGAGTCTGCTACTTTGACTTCAGT TCCATCAAGGTCTGTCATAGAAACACAAACAGTAAAACATGAAGGAAATATTAATGTCATCAGTGTTTCATTGAAGAAT ACCACAACAGAATATGAAGAGCACATAGGAACAGGTGGACCAGTGACATCAGTTTCAAGTACTGGATCAGACGGCTTGA CAGAGGAGTCTGAAGTAGCAATTGAGATGTCTGAGAATGTCTTTAGTACAGAAAACCAAGGAGAACCAACACAAGAAGC AGTACCAACTTACACAGCTCCTTCTGACATCAAATCAAGACTTGGTTCTAGAAGAGAGGTAACATCCCATGTTACACCA GTAATTAGAACAAAGGATTTAGAAACAGCTGAGGTCACATCATCTCCAGAATCAGTTGTGAATAACAGCACTCTTGACA CCATGGTGACACATGGTACCATAAGAGCAGTAGCTGAGTCTACAGAAAGCAAAAAGGGAAAAGGTAGCTTTTCAGCTGT TTCTTTGGGCAAAATTCTAATGATTGAGCATGGCTCTGGAGAAGAACTCAAAGTTGACAGCAGCACCACAAAATTAATG TCTAATGGACCTACGGAAAAACTGCTTGGAAGCCATTTTTCATTTTTTGACCAGGGATCTGGTGAAGCAGAAACTTTAA CTGAATCATTTACAAAAGCATCAGTTTCACCCACTGGGAAACCAGAACCACAGGAGCAGTATGGCAGAAAAACTGTCAG CATGCCATCCGCTGTGGTTCATGCCTACACTGCTGAACCTAATGAGCTAGTTACAAGTACTGAACATGACATAACCTCT TTGCAAACTGTAACTGATACAGAAATGGAAGAGAAAGCAGCTAATGAGCTCACAGTGACATCTTTTGCTACAAATCTTC CTTTGTCAGAAGATGTGCATTCATGGGAAGACAGGCCAAGGGAAATTTTACCAAAAGCAATAGAATCTTCTGGAGAAGC AACGGAAGATCCATTCTTCATAAGTACACAGGCTAACCATGAACATGTGGAATTTTTAAGTGTTCCCACTATCAGACCA CACTCAGAAGAAAACAAAGTAGAAGCTGAATCTGATGAAAAAATATTGTTGCCATTTAATAATGACAGAGTTACTGAGT CTGCAGTAATTGAAAGGAAATACTTAAGTTCTCCATTTACAGATACAGAGCAGGAGGAGGAGTTGGTACAAAATATTTT TCCTACAGAAGATATTCCCAGATTATTCCTGACACCTAAGGAAGAAAAACCAACAAACAATGAGCTCATCAGTGATCCA TTATTTTCAGGACAAGGATCAGGAGATGAATTTACTGTTATTCCTTCAGTAGAGTCATTGGCTGTGAAAGAAACCACAA ATACTTTAAGTCCTTGGCCTTTTCATCCTGCAAGTGTAGGACCTAAACTTTCCACTGACAAGACACAAGTCTTTGAAAG TGGAAGCACTGACTCAAATGCTGAAATTAATGAAGAAATTACAACTACTGCAGCAGAGCTGACAGAAACTGCATACTCA ATGGCCACCAGTTCCCCAGCTTTAGAAGAAGAGAGCTCCAGTCACTCCAACTCTAAAGACAAAGATATTACACATTATT TTCTTGTGATTGAAGACCCGTACAACAAGGAAATGGACCATAGAAGGGGAGAAAATGGAACGAGCAGGCCACTACCTAC TCCTGGAGATGTTTCTCTTGAGGAATCTTCACATATGCTGACTACAGATGACGTAACTCCTGTTTCTGTAATACTGAGC GAAACTCCTTATCTTGAAATGGGAAAATCTCTTGCAACATCAGCAACAAAAATGCCAAGCAGAGTATTACCTGAAAGTT CTGGAGAGGGATCTGGTTGGGATGGTGTTTCAGATTCATTTGCCCCTGATACATTGACTCATTCAACAGCACCTTCTGT GATGGAAGTTGAACTAACTGCTTCATCTCATATTCCTGGGGTTTATTCTGAAGTTATGACAACACATGTACCGGGAGAT GGATCACAGACTGTAATCACAGGATTAGCATCTCTTTTTACAGAAGAAAAAGAGATTGTGGCAAATCGAACTGCTGCTG ATCCAAAGACAGGTACATCGGAAGAACTGACGAGTGATACTGGGATGTCACTGGATATTATTCCAGTGGTTGATGATAG AAGACACGTTACACTGAATGTTTCTGTTTATGGTGATATTACACTAATTGAAGAACGACTTCAAATCCCATCAGAAAAA ACAACTATTATAGACATGGATCACTCCAAGTCAATGCCTGAAGATATAATAAGTGTGCAGACAATGCCAAATCTTGTCA TACGGTCAACACAAGTCAGTGATGATAACATGAAAGCTGAAGAGGACAAGTATGATTCAATACTTAATTTTTCTACAGT TGAAGAGAATTCATTTGGATCTGGTGATAATCTTTCTTTGACTACTTCCATTCAGCCAAGTAGTGAATCAGTAACAGCT GGACATGGACCAAAATTAGTAGACAAGGATTTAGGTTCAGGGTATGCCATGCAGTTTGCAACAGAAACTCTCACAACTA CTGTACTCAATGAATTGGGAATTTTCCTTCCAACAGTACCCTCACTGGTAAGCCCACATATGCCTCATGAGTCTAAAGA ATCTGAGTTCGAGGCTAAACACATTGGAAGGACAAGCACAACAGATGATGTGTATGAGCCTTATACTTCAGCAAACAAC CAAGTAATAACAGATCAAAGCAAAACAATGTCTATTTCTGGATTTTCTGGAATGGGCCAAGAAGAATCAGGTGATAAAA AGCCTATGATTCCTTCCCTTACACCAGATTTAACTATGGAGACAGAAAAGGCTTTGACAACTGACACATTTGATGTAAG TATGGTCACCACGCAGAGTATGTCCCAACATGCAACTGTATCATCTAGCAGTAGTGAAGAAAAGCATTCTACAGTATAC ATGCAAACAAAATCAGCATCAACAGAGTATGAAGAAACAGACTCAGTGAGTTTAAATTCTGTATCTCAAAATCCTAAAT CCTCAGTAACTGTTTGGTTAGTTAATGGAGTATCTAAATATCCTGAGGTAATTATTCCAAGTACATCATCTGCAAAGGA TTCAGATCAGTCTGATCATTCCTCAGATGGAACTTTCAAAGAGGTTAGTTCTGATATGGCAGCAACTTATAAACCACCC ACTACAGACCTAGATACAACAGTGTCTTCTCTGCTAGTGTTTTCTCCTGAGCCTGAATCAGAAAGCATATCTACTGAAA GTACTCCTCACTTTAATAAATTTGTAACGGAGAGGTCAGAGGAGACAGAATCTTCTGTAAATGATCTGATTATTGAGGA AAATGCTACTGTTTCTGGAGATTCTCCATCAATACATGACTACCCTACTGCTTTTTGGAATTTTGGAGAAAGAACAAGC ACAGATGTTCCAAAACTTAGCACAATAGAAGTTGAATTTTCTTCAGAGAGAGTGAAAAATCCCAGTCAAGAAAGTGATA GGAGTACTGAGAGAGAGAGACCAAGGCTTTCATCCGCACCTGTTTCAGATTCACCCAACAGTATAGAAGTTGGAGTATT TAAACCTGACCAGGAAGCAGTTACAATGCTAACTTCATCTTTGGAGCCTTTGGATAGAAGTTTGGAAACACAGTCAGCT TTGCTTGGTCCTTTGCTTGGTCAACAGGAGATCACAACAATTTCTTCTAACATTGCAACAAACAATACAGCCCCTGGAA ACAATCCTTATTCAAATGAACAGTCAACAATAAGCTCTGAGTTGCTAAATACTATTGAACTTGTAACTTCATCATTTTC CCTTCCGGAAGTCACTAATGGATCAGATTTCCTGATTGGCACAAGTGTGGGTTCGGTTGAAGGCACAGCAGTGCAAATT CCAGGACAGGATCCATGCAAAAGTAATCCCTGCCTTAATGGTGGTACCTGCTATCCACGTGGTTCATTTTACATCTGTA CATGTTTGCCAGGTTTCAATGGTGAGCAGTGTGAATTAGATATTGATGAATGCCAGTCTAACCCATGCCGCAATGGAGC CACATGTAT
Citation	Hudson KS, Andrews K, Early J, Mjaatvedt CH, Capehart AA. Versican G1 domain and V3 isoform overexpression results in increased chondrogenesis in the developing chick limb in ovo. Anat Rec (Hoboken). 2010 October; 293(10): 16691678.
Copyright	Copyright © 2010 Wiley-Liss, Inc.
Comments	The probe was amplified by PCR using the following primers: 5_-TACCTCAGACTGTCACAGATGG-3_ and 5_-ATACATGTGGCTCCATTGCGGCA-3_ and was extracted from NCBI Acc# NM_204787.1.

Stage

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stage 25

Wing
Wing Mesenchyme

Entry 2

GEISHA Id	VCAN.Singh.2016	Data Source	Publication
Complete cDNA Template Probe	show GCAGTATTAGCCCCTGAGCTCCACTTTTGGAACCATCGTTTGCTATTTCCTTGGAGAACAAAATTCCTGCTAAACAGGA TCATTTGTTTGCGGTTAGGTGGGTGAGATCTGCGTGACTTGCGGAAGAATCAGGATCTTGGCTTTCTAGAAAGAAGTGT GTGCACAAAGAGATTATAGGTTGTAATTGGTGCAGTGTGTGCGTAGGAGTTTTTCTGTTATGATTTCTTACCCTGGTCA GTGTGACAGCTGACCTGGATGTGTACAGTAAGTGTAGAAGGTAATGTTTGAAAGGACCTCTCCACATAAAGGACTGTGT CTGATTATTTCTGGGAAGTTCCCTGCACAACTCTGGGCACAGGAAAAAAAAGGACACAGCACAAAAGGGAAAATAAAGA ACAGTTAAGTTCCCTGGGATACACTTTCTAAAATAGCATTGCAAATACAGAGACAGATGAGAGGTTTATGCCTTTGTGT ATGTGGGGACGATACAAAAAGTTGTAAGGGACGAGGAGAAGAATAGAAAACACAGCCAACGTTTAAGAGTTTGTCCTAG GCAGAGCTACCCGTGAGGCAGCCTTGCCACGGCTTATCGGGAACACTGATTACAGTCCCAGACAATATCTCTTACAAAA TGGCAGTGACCGCTCCATCCTGGGTACCTGACCTTTATATGTTCAGCCTGGGCTGGTTAGCTGGTTCATTCCCCACCCC TGCCCCAAACAATGCACATAATACTTTCTCTATTATATTATCTGTAAAGGGTATATTGTGAAT
Citation	Singh PN, Ray A, Azad K, Bandyopadhyay A. A comprehensive mRNA expression analysis of developing chicken articular cartilage. Gene Expr Patterns. 2016 Jan;20(1):22-31.
Copyright	Copyright © 2015 Elsevier B.V. All rights reserved.
Comments	The complete cDNA template sequence was obtained from the information provided in the publication as described in Singh et al. 2016. The probe was synthesized from cDNA clone ChEST644d9.